Here are the pes footprints of the two giant ankylosaurs (not to scale) with my interpretations on the right:
Pula Albian (mid-Cretaceous) ankylosaur right pes lenght: 85-95 cm (A, B) - I am standing on it's toe #5 of it's four pes toes;
Tumbler Ridge (Canada) late Cretaceous anky has a scale of 10 cm for size comparison (about 44 cm total pes length) (E, F) Photo credit: Rich McCrea.
It is interesting that some members of Thyreophora - a subgroup of the ornithischian dinosaurs
which include well-known suborders Ankylosauria and Stegosauria, had diverse number of toes.
While some of the ankylosaur ichnogenera had three, the others had four toes on their hind feet.
Thus it is possible to miss identify the thyreophoran three-toed pes tracks for iguanodontoid ones and vice versa, especially if the front feet impressions are missing or are poorly preserved. Despite of the enormous record size of the Pula, Istrian tracks it is less likely that these were artifacts.
The Tumbler Ridge site
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